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Exchange Server 2003/2010 Coexistence Mail-flow Issues

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I've installed Exchange 2010 in a 2003 coexistence scenario. 2010 was deployed with CAS,HUB, and Mailbox roles. 

The installation went through smoothly, and the default RGC was created.

On the 2010 Server, when I create a new users with mailbox, that user is no able to send or receive mail from anywhere.

2010 user to 2010 user does not work
2003 user to 2010 user does not work

Mail delivery between 2003 users works fine. 

The RGC message queue in ESM 2003 shows that the messages are queued and keep retrying. 
I have deleted the RGC twice and created new ones, I have un-installed the HUB and CAS roles and re-installed them, and I have also re-run setup /preparead and setup /preparelegacyexchangepermissions

I'm not able to make any sense of the message tracking on the 2010 side. It just shows a bunch of entries: 

EventID: NOTIFYMAPI

Source: STOREDRIV 

When I use the best practices analyzer to test mailflow to a 2010 user from the outside, the test comes back with a pass even though the message itself isn't delivered to the specified mailbox. 

There is no smart host defined on the default SMTP virtual server on 2003.
There is no SMTP Send Connector for External or Internal mail configured on 2003 besides the 2010 RGC

Any ideas?

Here is the mail flow analyzer result:
     Testing inbound SMTP mail flow for domain 'user01@test.com'.
     Inbound SMTP mail flow was verified successfully.
     
     Additional Details
     
Elapsed Time: 3437 ms.
     
     Test Steps
     
     Attempting to retrieve DNS MX records for domain 'test.com'.
     One or more MX records were successfully retrieved from DNS.
     
     Additional Details
     
MX Records Host mail.test.com, Preference 0
, Host test.com, Preference 10
Elapsed Time: 110 ms.
     Testing Mail Exchanger mail.test.com.
     This Mail Exchanger was tested successfully.
     
     Additional Details
     
     Test Steps
     
     Attempting to resolve the host name mail.test.com in DNS.
     The host name resolved successfully.
     
     Additional Details
     Testing TCP port 25 on host mail.test.com to ensure it's listening and open.
     The port was opened successfully.
     
     Additional Details
     
Banner received: 220 EX2K3w2K3.test.net Microsoft ESMTP MAIL Service, Version: 6.0.3790.3959 ready at Wed, 5 Feb 2014 14:40:44 -0500
Elapsed Time: 368 ms.
     Analyzing SMTP Capabilities for server mail.test.com:25
     SMTP Capabilities were analyzed successfuly.
     
     Additional Details
     Attempting to send a test email message to user01@test.com using MX mail.test.com.
     The test email message was delivered successfully.
     
     Additional Details
     
Elapsed Time: 621 ms.
     Testing the MX mail.test.com for open relay by trying to relay to user Admin@TestExchangeConnectivity.com.
     The Open Relay test passed. This MX isn't an open relay.
     
     Additional Details
     Testing Mail Exchanger test.com.
     This Mail Exchanger was tested successfully.
     
     Additional Details
     
     Test Steps
     
     Attempting to resolve the host name test.com in DNS.
     The host name resolved successfully.
     
     Additional Details
     Testing TCP port 25 on host test.com to ensure it's listening and open.
     The port was opened successfully.
     
     Additional Details
     Analyzing SMTP Capabilities for server test.com:25
     SMTP Capabilities were analyzed successfuly.
     
     Additional Details
     Attempting to send a test email message to user01@test.com using MX test.com.
     The test email message was delivered successfully.
     
     Additional Details
     Testing the MX test.com for open relay by trying to relay to user Admin@TestExchangeConnectivity.com.
     The Open Relay test passed. This MX isn't an open relay.
     
     Additional Details

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